Ahead is another floor made of skull tiles with concealed spikes. In the co-op game, there are no grapple rings. Instead each pressure pad controls one of 3 wooden ledges above the spikes. So one player must depress the pads while another jumps across the ledges.
Start by having one character run out to the second pressure pad.
Then she can retrieve the red skull (4/5) and run carefully back to the first pressure pad.
(In these screenshots, the 2 characters then switch places, but it doesn't matter which character does what.) One character should continue to stand on the first pressure pad to keep the first wooden ledge raised, while the other jumps onto it. The character below should then run onto the second pressure pad to raise the next ledge.
As soon as the second ledge rises, the character above should jump onto it before the first ledge collapses.
Then the character below should run onto the third pressure pad.
As soon as the third ledge rises, the character above should jump onto it before the second ledge collapses. She can then jump over to the pressure pad on the upper level to open the gate below so the other character can exit.
The 4-player co-op game is basically the same as the 2-player, but instead of timing the jumps so carefully, all but one character can take turns moving across the spikes onto the pressure pads. While they hold up the ledges, one player can jump across and open the gate so the others can get out of the spikes. I'm not positive about the 3-player version, but I imagine it is similar.
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