Advance just far enough to kill the crocodiles in the pool on the left.
Then step out into the middle of the room where there are gems scattered on the floor. The gates ahead open and 2 of Sobek's minions emerge. Don't kill them yet! You'll need at least one of them alive for the next Challenge: Use the Trap Platforms to Dunk an Enemy.
To accomplish this, swim across the pool on the left. Then use the Staff of Osiris to raise a column underneath you, so you can reach the ledge above.
Stand near the Eye of Horus switch and wait for one of the crocodile men to leap up onto the wooden platform. Then shoot the switch...
...to collapse the platform, plunging the enemy into the water and completing the challenge. Your reward is a MAX AMMO UPGRADE.
Naturally a little water doesn't hurt Sobek's warriors. The guy you just dunked jumps back up and comes after you, along with his friend if you didn't already kill him. So be ready to finish them off with weapons and bombs.
As far as I know, this challenge works the same way in co-op mode, and everyone gets the reward when you beat it.
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